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ethology zoology animal behaviour animal science sensory receptor neuron research methods ug course survivorship mortality natality immigration emigration density shelford's laws of tolerance liebig law of minimum earnst haeckel wallace blackman law of limiting factor father of ecology mac arthur odum voltera lotka darwin linnaeus database data non probability sampling random sampling cluster sampling sampling research methodlogy epidermis nerves nervous system nerve senses kornard lorenz animal behavior stimulus sensitization method learning behaviour conditioned learning ecology response stimuli