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stable isotopes oxygen isotopes earthworms food web calcite mid-cayman rise rimicaris hybisae diet hydrothermal vents rhine meuse europa freshwater mussels origin of life evolution biology freshwater bivalves life uric acid urea moon meteorite solar system comet asteroid granules climate calcium carbonate carbon hydrothermal ammonia stenohaline euryhaline hypoosmotic hyperosmotic osmoconformer