Discomycetes are a large group of ascomycete fungi that produce fruiting bodies called apothecia. They can be saprobes, parasites, or form lichens and mycorrhizal relationships. Discomycetes are classified based on whether their apothecia have an operculum or not. Some economically important discomycetes include edible mushrooms in the order Pezizales and truffles in the order Tuberales. The apothecium contains the hymenium layer where asci and ascospores are produced.