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genetics plant breeding qtl mapping drought marker assisted selection wheat molecular markers drought tolerance plant genetics root dna markers root angle aflp brassica oleracea pcr dna brassica napus genetic genetic diversity genetics punica granatum p. prot gwas glucosinolates faba bean frost tolerance gene annotation barley salt tolerance proline physiological traits leaf wilting tissue culture transformation balanites aegyptiaca in vitro propagation gynogenesis maize pollen genetic diversity potyviruses cucurbit qtl