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§ª§à§ß§ß§í§Ö §ä§à§Ü§Ú §Ó §á§Ö§Û§ã§Þ§Ö§Ü§Ö§â§ß§í§ç §ß§Ö§Û§â§à§ß§Ñ§ç §£§Ö§Õ§å§ë§Ñ§ñ §â§à§Ý§î-§Ò§í§ã§ä§â§à §Ñ§Ü§ä§Ú§Ó§Ú§â§å§Ö§Þ§í§Û §Ú §Þ§Ö§Õ§Ý§Ö§ß§ß§à §Ú§ß§Ñ§Ü§ä§Ú§Ó§Ú§â§å§Ö§Þ§í§Û §Ó§ç§à§Õ§ñ§ë§Ú§Û I Na+ §¡§Ü§ä§Ú§Ó§Ñ§è§Ú§ñ: -60 §Þ§£. §±§Ú§Ü: -30 §Þ§£. I L - §á§Ñ§ã§ã§Ú§Ó§ß§í§Û §ä§à§Ü §å§ä§Ö§é§Ü§Ú, §Ó §à§ã§ß§à§Ó§ß§à§Þ, §¬ + §°§ä§ß§à§ê§Ö§ß§Ú§Ö gNa + /gL §Ò§à§Ý§î§ê§Ö §ä§Ñ§Ü§à§Ó§à§Ô§à §Ó §ß§Ö§á§Ö§Û§ã§Þ§Ö§Ü§Ö§â§ß§í§ç §Ü§Ý§Ö§ä§Ü§Ñ§ç Butera, Smith. 1999
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crasgmu
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crasgmu
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crasgmu
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crasgmu
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crasgmu
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crasgmu
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  • 1. §³§à§Ó§â§Ö§Þ§Ö§ß§ß§í§Ö §á§â§Ö§Õ§ã§ä§Ñ§Ó§Ý§Ö§ß§Ú§ñ §à §ã§ä§â§å§Ü§ä§å§â§Ö §Õ§í§ç§Ñ§ä§Ö§Ý§î§ß§à§Ô§à §è§Ö§ß§ä§â§Ñ
  • 2. §¥§í§ç§Ñ§ä§Ö§Ý§î§ß§í§Û §è§Ö§ß§ä§â §Ó medulla oblongata §³§á§Ú§ß§Ñ§Ý§î§ß§í§Ö §Þ§à§ä§à§ß§Ö§Û§â§à§ß§í §¥§í§ç§Ñ§ä§Ö§Ý§î§ß§Ñ§ñ §Þ§å§ã§Ü§å§Ý§Ñ§ä§å§â§Ñ §¥§í§ç§Ñ§ß§Ú§Ö¨C §á§â§à§è§Ö§ã§ã §Ô§Ñ§Ù§à§à§Ò§Þ§Ö§ß§Ñ §Þ§Ö§Ø§Õ§å §Ü§Ý§Ö§ä§Ü§Ñ§Þ§Ú §Ú §à§Ü§â§å§Ø§Ñ§ð§ë§Ö§Û §ã§â§Ö§Õ§à§Û
  • 3. §¡§ß§Ñ§ä§à§Þ§Ú§ñ §Õ§í§ç§Ñ§ä§Ö§Ý§î§ß§à§Ô§à §è§Ö§ß§ä§â§Ñ Nucleus ambiguus Nucleus solitarius I §á§â§Ö-§¢§Ö§ä§è§Ú§ß§Ô§Ö§â§Ñ §Ü§à§Þ§á§Ý§Ö§Ü§ã E E I §¢§Ö§ä§è§Ú§ß§Ô§Ö§â§Ñ §Ü§à§Þ§á§Ý§Ö§Ü§ã
  • 4. §´§Ú§á§í §Ñ§Ü§ä§Ú§Ó§ß§à§ã§ä§Ú §Õ§í§ç§Ñ§ä§Ö§Ý§î§ß§í§ç §ß§Ö§Û§â§à§ß§à§Ó §ª§ß§ã§á§Ú§â§Ñ§ä§à§â§ß§í§Ö §â§Ñ§ß§ß§Ú§Ö §ª§ß§ã§á§Ú§â§Ñ§ä§à§â§ß§í§Ö §á§à§Ù§Õ§ß§Ú§Ö §ª§ß§ã§á§Ú§â§Ñ§ä§à§â§ß§í§Ö §á§à§Ý§ß§í§Ö §¿§Ü§ã§á§Ú§â§Ñ§ä§à§â§ß§í§Ö §â§Ñ§ß§ß§Ú§Ö §¿§Ü§ã§á§Ú§â§Ñ§ä§à§â§ß§í§Ö §á§à§Ù§Õ§ß§Ú§Ö §¿§Ü§ã§á§Ú§â§Ñ§ä§à§â§ß§í§Ö §á§à§Ý§ß§í§Ö §ª§ß§ã§á§Ú§â§Ñ§ä§à§â§ß§à-§ï§Ü§ã§á§Ú§â§Ñ§ä§à§â§ß§í§Ö §¿§Ü§ã§á§Ú§â§Ñ§ä§à§â§ß§à-§Ú§ß§ã§á§Ú§â§Ñ§ä§à§â§ß§í§Ö
  • 5. §¤§â§å§á§á§í §Õ§í§ç§Ñ§ä§Ö§Ý§î§ß§í§ç §ß§Ö§Û§â§à§ß§à§Ó §¤§Ö§ß§Ö§â§Ñ§ä§à§â§í §Õ§í§ç§Ñ§ä§Ö§Ý§î§ß§à§Ô§à §â§Ú§ä§Þ§Ñ §¶§à§â§Þ§Ú§â§å§ð§ë§Ú§Ö §Õ§í§ç§Ñ§ä§Ö§Ý§î§ß§í§Û §á§Ñ§ä§ä§Ö§â§ß §â§Ñ§ß§ß§Ú§Ö §Ú§ß§ã§á§Ú§â§Ñ§ä§à§â§ß§í§Ö §á§à§ã§ä§Ú§ß§ã§á§Ú§â§Ñ§ä§à§â§ß§í§Ö §á§à§Ý§ß§í§Ö §Ú§ß§ã§á§Ú§â§Ñ§ä§à§â§ß§í§Ö §á§à§Ù§Õ§ß§Ú§Ö §Ú§ß§ã§á§Ú§â§Ñ§ä§à§â§ß§í§Ö §ï§Ü§ã§á§Ú§â§Ñ§ä§à§â§ß§í§Ö D. W. Richter, 1982,1992
  • 6. §®§à§Õ§Ö§Ý§Ú §Õ§í§ç§Ñ§ä§Ö§Ý§î§ß§à§Ô§à §â§Ú§ä§Þ§à§Ô§Ö§ß§Ö§Ù§Ñ: 1. Network model §ã§Ú§ß§Ñ§á§ä§Ú§é§Ö§ã§Ü§Ñ§ñ §â§Ö§Ô§å§Ý§ñ§è§Ú§ñ §Ó§Ö§Õ§å§ë§Ñ§ñ §â§à§Ý§î §ä§à§â§Þ§à§Ù§ß§í§ç §ã§Ó§ñ§Ù§Ö§Û §Ó§í§ã§à§Ü§Ñ§ñ §Ù§ß§Ñ§é§Ú§Þ§à§ã§ä§î §â§Ö§Ô§å§Ý§Ú§â§å§ð§ë§Ú§ç §æ§Ñ§Ü§ä§à§â§à§Ó (§ç§Ö§Þ§à- §Ú §Þ§Ö§ç§Ñ§ß§à§â§Ö§è§Ö§á§ä§à§â§í) §±§â§Ú§é§Ú§ß§Ñ §Ñ§Ó§ä§à§Þ§Ñ§ä§Ú§Ú §¥§¸: §ß§Ñ§Ý§Ú§é§Ú§Ö §â§Ñ§Ù§Ý§Ú§é§ß§í§ç §ä§Ú§á§à§Ó §ß§Ö§Û§â§à§ß§à§Ó, §à§Ò§ì§Ö§Õ§Ú§ß§Ö§ß§ß§í§ç §à§á§â§Ö§Õ§Ö§Ý§Ö§ß§ß§à§Û §ã§ä§â§å§Ü§ä§å§â§à§Û §ã§Ó§ñ§Ù§Ö§Û
  • 7. 4 2 1 3 5 6 §¥ §® Network model
  • 8. §®§à§Õ§Ö§Ý§Ú §Õ§í§ç§Ñ§ä§Ö§Ý§î§ß§à§Ô§à §â§Ú§ä§Þ§à§Ô§Ö§ß§Ö§Ù§Ñ: 2. Pacemaker-network model §Ô§Ö§ß§Ö§â§Ñ§è§Ú§ñ §Õ§í§ç§Ñ§ä§Ö§Ý§î§ß§à§Ô§à §â§Ú§ä§Þ§Ñ-§ã§á§à§ß§ä§Ñ§ß§ß§í§Û §ß§Ö§Û§â§à§ß§ß§í§Û §á§â§à§è§Ö§ã§ã §á§Ö§Û§ã§Þ§Ö§Ü§Ö§â§ß§í§Ö §á§â§Ö§Ú§ß§ã§á§Ú§â§Ñ§ä§à§â§ß§í§Ö §ß§Ö§Û§â§à§ß§í §â§Ñ§ã§á§à§Ý§à§Ø§Ö§ß§í §Ó §á§â§Ö§Õ§Ö§Ý§Ñ§ç §Ü§à§Þ§á§Ý§Ö§Ü§ã§Ñ §á§â§Ö-§¢§Ö§ä§è§Ú§ß§Ô§Ö§â§Ñ §â§Ú§ä§Þ§à§Ô§Ö§ß§Ö§â§Ú§â§å§ð§ë§Ñ§ñ §à§ã§ß§à§Ó§Ñ §Ó§ã§ä§â§à§Ö§ß§Ñ §Ó §Ü§à§Þ§á§Ý§Ö§Ü§ã§ß§å§ð §ß§Ö§Û§â§à§ß§ß§å§ð §ã§Ö§ä§î §Õ§Ú§ß§Ñ§Þ§Ú§é§Ö§ã§Ü§Ú§Ö §Ó§Ù§Ñ§Ú§Þ§à§Õ§Ö§Û§ã§ä§Ó§Ú§ñ §Þ§Ö§Ø§Õ§å §ã§Ú§ß§Ñ§á§ä§Ú§é§Ö§ã§Ü§Ú§Þ§Ú §ã§Ú§Ô§ß§Ñ§Ý§Ñ§Þ§Ú §Ú §Ó§ß§å§ä§â§Ö§ß§ß§Ú§Þ§Ú §ã§Ó§à§Û§ã§ä§Ó§Ñ§Þ§Ú §ã§Ñ§Þ§à§Û §Ü§Ý§Ö§ä§Ü§Ú Smith et al, 1995, 1999
  • 9. §ª§à§ß§ß§í§Ö §ä§à§Ü§Ú §Ó §á§Ö§Û§ã§Þ§Ö§Ü§Ö§â§ß§í§ç §ß§Ö§Û§â§à§ß§Ñ§ç §£§Ö§Õ§å§ë§Ñ§ñ §â§à§Ý§î-§Ò§í§ã§ä§â§à §Ñ§Ü§ä§Ú§Ó§Ú§â§å§Ö§Þ§í§Û §Ú §Þ§Ö§Õ§Ý§Ö§ß§ß§à §Ú§ß§Ñ§Ü§ä§Ú§Ó§Ú§â§å§Ö§Þ§í§Û §Ó§ç§à§Õ§ñ§ë§Ú§Û I Na+ §¡§Ü§ä§Ú§Ó§Ñ§è§Ú§ñ: -60 §Þ§£. §±§Ú§Ü: -30 §Þ§£. I L - §á§Ñ§ã§ã§Ú§Ó§ß§í§Û §ä§à§Ü §å§ä§Ö§é§Ü§Ú, §Ó §à§ã§ß§à§Ó§ß§à§Þ, §¬ + §°§ä§ß§à§ê§Ö§ß§Ú§Ö gNa + /gL §Ò§à§Ý§î§ê§Ö §ä§Ñ§Ü§à§Ó§à§Ô§à §Ó §ß§Ö§á§Ö§Û§ã§Þ§Ö§Ü§Ö§â§ß§í§ç §Ü§Ý§Ö§ä§Ü§Ñ§ç Butera, Smith. 1999
  • 10. §³§Ú§ß§Ñ§á§ä§Ú§é§Ö§ã§Ü§Ú§Ö §Ó§Ý§Ú§ñ§ß§Ú§ñ §ß§Ñ §ß§Ö§Û§â§à§ß§í 1. §¤§¡§®§¬ (§¡) §â§Ö§è§Ö§á§ä§à§â§í ¨C §Ñ§Ü§ä§Ú§Ó§Ú§â§å§ð§ä Cl - §Ü§Ñ§ß§Ñ§Ý§í 2. §¤§¡§®§¬ (§£) §â§Ö§è§Ö§á§ä§à§â§í ¨C §Ñ§Ü§ä§Ú§Ó§Ú§â§å§ð§ä §¬ + §Ü§Ñ§ß§Ñ§Ý§í §ä§à§â§Þ§à§Ø§Ö§ß§Ú§Ö §ï§Ü§ã§á§Ú§â§Ñ§ä§à§â§ß§í§ç §ß§Ö§Û§â§à§ß§à§Ó §Ó§à §Ó§â§Ö§Þ§ñ §á§à§ã§ä§Ú§ß§ã§á§Ú§â§Ñ§ä§à§â§ß§à§Û §æ§Ñ§Ù§í c §Ö§Ý§Ö§Ü§ä§Ú§Ó§ß§í§Û §Ñ§ß§ä§Ñ§Ô§à§ß§Ú§ã§ä CGP55845A 3. P §Ö§è§Ö§á§ä§à§â§í §Ü §Ô§Ý§Ú§è§Ú§ß§å¨C §ä§à§â§Þ§à§Ø§Ö§ß§Ú§Ö §á§à§Ù§Õ§ß§Ú§ç §Ú§ß§ã§á§Ú§â§Ñ§ä§à§â§ß§í§ç §ß§Ö§Û§â§à§ß§à§Ó §â§Ñ§ß§ß§Ú§Þ§Ú §ï§Ü§ã§á§Ú§â§Ñ§ä§à§â§ß§Ñ§Þ§Ú §ß§Ö§Û§â§à§ß§Ñ§Þ§Ú W. Zhang et al. 2002
  • 11. §ª§ß§ß§Ö§â§Ó§Ñ§è§Ú§ñ §Õ§í§ç§Ñ§ä§Ö§Ý§î§ß§à§Ô§à §è§Ö§ß§ä§â§Ñ §±§ã§Ú§ç§Ú§é§Ö§ã§Ü§Ú§Ö §á§â§à§è§Ö§ã§ã§í §·§Ö§Þ§à§â§Ö§è§Ö§á§ä§à§â§í §®§Ö§ç§Ñ§ß§à§â§Ö§è§Ö§á§ä§à§â§í §´§Ö§â§Þ§à§â§Ö§è§Ö§á§ä§à§â§í §²§Ö§è§Ö§á§ä§à§â§í §Ó§Ö§â§ç§ß§Ú§ç §Õ§í§ç§Ñ§ä§Ö§Ý§î§ß§í§ç §á§å§ä§Ö§Û §¢§à§Ý§Ö§Ó§í§Ö §Ó§à§Ù§Õ§Ö§Û§ã§ä§Ó§Ú§ñ §°§â§Ô§Ñ§ß§í §é§å§Ó§ã§ä§Ó §®§Ö§Õ§Ú§Ñ§ä§à§â§í, §¢§¡§± §¥§® §¥ §¸
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